Elm

Elms are deciduous and semi-deciduous trees comprising the genus Ulmus in the family Ulmaceae. They are distributed over most of the Northern Hemisphere, inhabiting the temperate and tropical-montane regions of North America and Eurasia, presently ranging southward in the Middle East to Lebanon and Israel, and across the Equator in the Far East into Indonesia.

Elms are components of many kinds of natural forests. Moreover, during the 19th and early 20th centuries, many species and cultivars were also planted as ornamental street, garden, and park trees in Europe, North America, and parts of the Southern Hemisphere, notably Australasia. Some individual elms reached great size and age. However, in recent decades, most mature elms of European or North American origin have died from Dutch elm disease, caused by a microfungus dispersed by bark beetles. In response, disease-resistant cultivars have been developed, capable of restoring the elm to forestry and landscaping.

Description

The genus is hermaphroditic, having apetalous perfect flowers which are wind-pollinated. Elms leaves are alternate, with simple, single- or, most commonly, doubly serrate margins, usually asymmetric at the base and acuminate at the apex. The fruit is a round wind-dispersed samara flushed with chlorophyll, facilitating photosynthesis before the leaves emerge. The samarae are very light, those of British elms numbering around 50,000 to the pound (454 g). (Very rarely anomalous samarae occur with more than two wings.) All species are tolerant of a wide range of soils and pH levels but, with few exceptions, demand good drainage. The elm tree can grow to great height, the American elm in excess of 30 metres (98 feet), often with a forked trunk creating a vase profile.

Taxonomy

There are about 30 to 40 species of Ulmus; the ambiguity in number results from difficulty in delineating species, owing to the ease of hybridization between them and the development of local seed-sterile vegetatively propagated microspecies in some areas, mainly in the Ulmus field elms (Ulmus minor) group. Oliver Rackham describes Ulmus as the most critical genus in the entire British flora, adding that ‘species and varieties are a distinction in the human mind rather than a measured degree of genetic variation’. Eight species are endemic to North America and three to Europe, but the greatest diversity is in Asia with approximately two dozen species. The oldest fossils of Ulmus are leaves dating Paleocene, found across the Northern Hemisphere.

The classification adopted in the List of elm species is largely based on that established by Brummitt. A large number of synonyms have accumulated over the last three centuries; their currently accepted names can be found in the list of Elm synonyms and accepted names.

Botanists who study elms and argue over elm identification and classification are called “pteleologists”, from the Greek πτελέα (elm).

As part of the suborder urticalean rosids, they are distantly related to cannabis, mulberries, figs, hops, and nettles.

Elm – Ecology

Elm propagation methods vary according to elm type and location, and the plantsman’s needs. Native species may be propagated by seed. In their natural setting, native species, such as wych elm and European white elm in central and northern Europe and field elm in southern Europe, set viable seed in “favourable” seasons. Optimal conditions occur after a late warm spring. After pollination, seeds of spring-flowering elms ripen and fall at the start of summer (June); they remain viable for only a few days. They are planted in sandy potting soil at a depth of 1 cm, and germinate in three weeks. Slow-germinating American elm will remain dormant until the second season. Seeds from autumn-flowering elms ripen in the fall and germinate in the spring. Since elms may hybridize within and between species, seed propagation entails a hybridisation risk. In unfavourable seasons, elm seeds are usually sterile. Elms outside their natural range, such as English elm U. minor ‘Atinia’, and elms unable to pollinate because pollen sources are genetically identical, are sterile and are propagated by vegetative reproduction. Vegetative reproduction is also used to produce genetically identical elms (clones). Methods include the winter transplanting of root suckers; taking hardwood cuttings from vigorous one-year-old shoots in late winter, taking root cuttings in early spring; taking softwood cuttings in early summer; grafting; ground and air layering; and micropropagation. A bottom heat of 18 °C and humid conditions are maintained for hard- and softwood cuttings. The transplanting of root suckers remains the easiest and most common propagation method for European field elm and its hybrids. For specimen urban elms, grafting to wych-elm rootstock may be used to eliminate suckering or to ensure stronger root growth. The mutant-elm cultivars are usually grafted, the “weeping” elms ‘Camperdown’ and ‘Horizontalis’ at 2–3 m (7–10 ft), the dwarf cultivars ‘Nana’ and ‘Jacqueline Hillier’ at ground level. Since the Siberian elm is drought tolerant, in dry countries, new varieties of elm are often root-grafted onto this species.[

Source:  WIKIPEDIA